See below for research with undergraduates at Amherst College.  Link to research publications and presentations at meetings.


Amherst College, Department of Biology Undergraduate Honors Theses


Atanga-McCormick, L. In progress, 2011-2012. Phylogeography of the widespread species, Lycium carolinianum, and the evolution of mating systems in mainland and island populations.


Keyes, T. J.  In progress, 2011-2012.  Lycium minimum (Solanaceae), an endemic Galapagos shrub... Phylogeography and characterization of its sexual system


O’Keeffe, K. In progress, 2011-2012. Evolution of mating system genes in self-incompatible and self-incompatible populations of Lycium carolinianum.


Kamath, A. 2011. The evolution of floral morphology accompanying the transition from cosexuality to gender dimorphism in Lycium californicum (Solanaceae). B.A. Thesis, Department of Biology, Amherst College, U.S.A.


Beck, T. R. 2010. Phylogeography of Lycium californicum: intraspecific variation in sexual system and ploidy level. B.A. thesis, Department of Biology, Amherst College, U.S.A.


Damashek, J. 2009. Uncovering a tale of worldwide plant dispersal: evolutionary relationships

of Chinese Lycium (Solanaceae). B.A. thesis, Department of Biology, Amherst College, U.S.A.


Dhond, M. 2009. Confirmation of a genetic bottleneck at the S-RNase locus in Old World

Lycium (Solanaceae). B.A. thesis, Department of Biology, Amherst College, U.S.A.


Barmettler, S. 2008. Phylogeography of Lycium californicum (Solanaceae) in the southwestern United States and Mexico. B.A. thesis, Department of Biology, Amherst College, U.S.A.


Whelan, A. P. 2008. The utility of nuclear conserved orthologous set II (COSII) markers for

species-level phylogenetic resolution in Lycium (Solanaceae). B.A. thesis, Department of Biology, Amherst College, U.S.A.


de Waal, C. 2007. Breeding Systems in African Lycium (Solanaceae). University of the Free State, Bloemfontein, South Africa (co-advised with Dr. Andor Venter)


Leffler, E. M. 2006. Ploidy level and self-incompatibility in five Lycium species (Solanaceae):

evidence from the S-RNase locus. B.A. thesis, Department of Biology, Amherst College, U.S.A.


Shak, J. R. 2006. Phylogenetic relationships of Old World Lycium (Solanaceae): Reticulate evolution in the African taxa. B.A. thesis, Department of Biology, Amherst College, U.S.A.


Blanton, J. M. 2005. Species-Level Phylogenetics for an American clade of Lycium (Solanaceae): exploring the utility of the genetic marker nitrate reductase. B.A. thesis, Department of Biology, Amherst College, U.S.A.


Neymark, A. 2005. Characterization of the S-RNase mating gene in polyploid Lycium exsertum. Department of Biology, Amherst College, U.S.A.


Savage, A. E. 2004. Molecular characterization of the S-locus in Lycium: breakdown of gametophytic self-incompatibility and the evolution of gender dimorphism. B.A. thesis, Department of Biology, Amherst College, U.S.A.


Amherst College, Summer 10-week Research Projects


2009. Cara J. Giaimo, Chloroplast sequence variation and plant barcoding


  1. 2008.Ambika Kamath, S-RNase diversity in Chinese Lycium


2008. Carolyn Whiting (Smith College), Does phylogenetic evidence support three species in Grabowskia?


2007. Jamie L. Kostyun, Allele specific amplification of the self-incompatibility gene (S-RNase)

in two species of Lycium (Solanaceae)


2007. Wen Zhang, Variation in noncoding chloroplast regions of Lycium


2006. Sara Barmettler, Development of a digital on-line database for genomic information


2006. Mukta Dhond, Lycium cestroides: extraction and isolation of the S-RNase gene controlling gametophytic self-incompatibility


2005. Alan C. Kwan, Lycium phylogenies: a study of the utility of nuclear and chloroplast

gene regions


2004. Denise Twum, The utility of chloroplast and nuclear gene regions for inferring phylogenetic relationships among Lycium (Solanaceae)


2003. Allison R. Schroder, The biogeography of Lycium (Wolfberry) based upon two chloroplast and one nuclear gene region